Before getting into the actual issue of invasive species, it is necessary to explain some basic terminology.
Alien species
A species that occurs outside its native (natural) range. This is the broadest term, encompassing all species that have entered an area through human activity or by natural means. It is the opposite of a native (indigenous) species that evolved in our area or arrived here by natural dispersal after the last ice age. A non-native species may arrive in a new range by introduction, by introduction or by natural means.
Adventive species
This term describes how the insect got here. Introduction occurs unintentionally - typically as a 'stowaway' in international trade. The insects are most often transported with timber, seedlings, fruit or in parcel or container shipments.
Introduced species
Although 'introduced' and 'introduced' are often used interchangeably in common parlance, in a more strict sense, introduction is the deliberate introduction by humans (e.g. for biological conservation, breeding, scientific research or as an ornamental species). However, when referring to 'introduced species' in general, it means all species that have crossed a geographical barrier thanks to humans.
A species that spreads naturally
Here we distinguish the concept of active migration, where the insect's range is gradually extended by direct flight of adults. This is usually a slower process that is limited by ecological barriers. Then there is passive migration, in which insects are transported by wind (especially small insects, e.g. aphids, thrips), by watercourses (larvae, pupae, drifting material) or by other animals (phoretic transmission on birds, mammals or other insects). Passive migration is usually of local or regional importance.
Invasive species
This is the narrowest and most alarming category. In order for a species to be considered invasive, it must meet three conditions:
- It is non-native to the area.
- It spreads uncontrolled and establishes viable populations in the wild.
- It causes damage - ecological (displacing native species), economic (destroying crops) or health.
Invasive species can be classified according to different aspects. In practice, the most commonly used are:
- invasion phase,
- degree of impact,
- the type of environment they affect.
✔️ Classification by phase of invasion
This division describes what stage of spread the species is in.
- Introduced species
A species has appeared in an area but has not established a permanent population. Individuals occur only irregularly, and cannot be sustained for long periods without further introduction from outside. They often survive only for a short time. This is a potentially invasive species that poses a risk for the future. - Naturalised (established) species
The species has established a stable, self-sustaining population but remains locally restricted. It does not disperse significantly into the surrounding area, its distribution is stable and it does not cause major damage. It has become part of the local fauna but is not aggressive. It is a so-called early invasive species for which intervention to limit its spread is still possible. - Invasive (expansive) species
A species that actively and rapidly spreads to new locations, increases its territory and causes demonstrable damage - displacing native species, altering communities, damaging economically important crops or woody plants. This is the highest and most problematic level. These are the species that can no longer be prevented. The aim of conservation is no longer eradication, but limiting the impact.
✔️ Classification by level and type of impact
This breakdown focuses on what the invasive species is doing harm.
- Ecologically significant invasive species
Species that threaten our biodiversity. They either directly eat our native species or take away their food and habitat (competition). - Economically important invasive species
Species that cause billions of dollars of damage to agriculture, forestry or warehouses. - Hygienically or medically important invasive species
Species that threaten human or animal health (transmit diseases, cause allergies). This is less common in beetles than e.g. mosquitoes, but there are species that cause skin irritation (e.g. various synanthropic species).
✔️ Classification by type of environment
This division allows a clear ordering of species.
- Invasive synanthropic species
These are tied to human dwellings, warehouses or industrial sites. Often survive in the long term only thanks to humans. - Invasive species of forest ecosystems
Invade trees, spreading in both economic and natural forests. They can have a major ecological and economic impact. - Invasive species of open landscapes and agriculture
Found in fields, meadows and orchards. Often have multiple generations per year = high reproductive potential. - Invasive species of aquatic and wetland ecosystems
These species are less common in beetles. However, when they do occur, they colonise very ecologically sensitive environments.
The following paragraphs explain the individual columns in the table below. The data sources used to create the table can be found in the menu "About project" -> "Sources".
The list has been compared with the family catalogues available on this website. The names of families and taxa in the list have been updated according to these catalogues. The names were taken as of 24 February 2026. The origin status was taken from the DAISIE database. Taxa displayed that were not listed in the DAISIE database at the time the list was prepared were added from other sources, if the information was known. Year of first occurrence of the taxon in the Czech Republic, if known. The field value can take one of the following values: The place from which the species was introduced to the Czech Republic. In addition to the basic regions listed above, abbreviations are added to further specify the region: The manner in which a non-native species entered a new area of distribution. Depending on the duration and nature of their occurrence, alien species are divided into the following categories: Eusynanthropic species – S, which survive only for a short time after introduction and reproduce only inside buildings, greenhouses and other heated structures, gradually disappearing or being destroyed (as indicated by the letter in brackets), or forming long-term populations in such places and possibly being dispersed by humans (partly spontaneously during the warm season); exceptionally, some of these species may form short-term populations in open spaces – SC. Accidentally non-native species – C, which, after escaping or being released into the open, form only short-term populations and then disappear again (this group does not include individuals that have escaped from captivity or individual captures of introduced animals that did not reproduce further). Naturalised, non-invasive species – N, which form long-term populations in the external environment, colonise suitable habitats at the site of introduction, do not spread, or if they do, only in the immediate vicinity of this site. Naturalised, post-invasive species (Pyšek et al., 2002) colonised this area in the past and have been known since the beginning of their study as fully naturalised, with more or less no changes in their distribution – *N, e.g. Aceria tristriata, A. erinea, Caloptilia roscipennella, Chromaphis juglandicola and Panaphis juglandis (all trophically linked to Juglans regia). These species evidently invaded Central Europe before 1900, and there is no way to determine whether they were dispersed with the host plant, followed it immediately, or invaded the area with a certain time lag. It appears that the invasions of many species occurred long after their host species had been introduced, which also suggests that in several cases, the dispersal of the host may not have been the only factor that initiated the invasion. For example, the invasion of the moth of the genus Gracillariidae, Phyllonorycter platani, did not occur until the 20th century (Šefrová, 2001), and the invasion of the bug Lygaeidae, Arocatus longiceps, did not occur until the end of the century (Stehlík & Hradil, 2000), although plane trees (Platanus spp.), their host species, had been planted in Central Europe for about 200 years. These species are considered invasive and are classified in another group. A somewhat special group consists of several species of fish and birds that do not reproduce naturally but are often kept in open spaces, sometimes in large numbers, and can thus significantly influence processes in semi-natural and natural ecosystems, as well as their biodiversity – M. Invasive species – I, representing that part of naturalised species that does not remain in the place of introduction but disperses from it and colonises areas of varying sizes. The speed of such animal invasions is sometimes significantly higher than that of plant species (see Richardson et al., 2000) and usually reaches kilometres or even hundreds of kilometres per year. For example, the current invasion of Cameraria ohridella in Europe is spreading at a rate of approximately 50 km per year (Šefrová, 2003). The status of non-native species and their occurrence is clearly shown in the overview table: The occurrence of naturalised species is limited to urban habitats (U) to which they are bound in terms of climate, environmental structure or food (e.g. they feed on ornamental plants, domestic animals and other synanthropic species), or spread across the landscape by occupying man-made habitats (agricultural crops, purpose-built vegetation, artificial water habitats, ruderal areas) (R), or penetrate semi-natural and natural communities (A). Animal and some plant parasites are closely linked to their hosts; their occurrence depends more on the occurrence of the host than on the nature of the habitat (H). The second letter added to the code specifies the habitat: The main entry in this field characterises the main type of nutrition (trophic strategy) of the species in question. The second letter in brackets specifies the potential impact, i.e. the possible economic or ecological significance of the species:
Taxon
Origin status (DAISIE)
Status
Name
Definition
A
Non-native
Non-native species from areas outside Europe.
C
Unclear origin
Species for which it is unclear whether they are native or introduced.
E
Transferred
A European species that has become alien outside its original range.
?
Unknown
Unknown origin status. Appears where there is insufficient information to determine status.
First occurrence in the Czech Republic
Place of origin of the taxon
Code
Region
AF
Africa
AM
America
AS
Asia
AUS
Australia
EU
Europe
MED
Mediterranean region
NZE
New Zealand
Code
Location specification
C
Central
E
Eastern
N
Northern
S
South
W
Western
Method of introduction
Code
Definition
A
Accidentally introduced species (Adventive species)
D
Introduced species, i.e. intentionally introduced (Introduced species)
S
Spontaneously introduced, i.e. a species spreading naturally
Invasive status
Code
Definition
C
occasional
I
invasive
M
reproduction (in open spaces)
N
naturalised, non-invasive
*N
naturalised, post-invasive
S
only eusynanthropic
(S)
occasional eusynanthropic occurrence (short-term occurrence after introduction)
Code
Definition
A
semi-natural or natural
H
occurring in the host
R
cultivated landscape (agricultural habitat, ornamental crops, ruderal plants, etc.)
U
settlements (urban habitat, relationship to urban environment, ornamental plants or domestic animals)
Food requirements
Code
Definition
Pa
zooparasitic / parasitoid
The species parasitises an animal host. Parasitoid means that larval development takes place on/inside the host and leads to its death (typical, for example, in some Staphylinidae or Cleridae).
Ph
phytophagous (often rather phytophagous)
Feeds on plants or parts of plants. This can include leaves, seeds, roots, flowers, etc.
Pl
planktophagous
Feeds on plankton (rather rare among beetles; typically larvae of aquatic species).
Po
polyphagous
A species with no dietary specialisation; utilises a wide range of sources (e.g. various types of organic matter).
Pr
predator
Actively hunts other animals.
Sa
saprophagous
Feeds on dead organic matter (detritus, decomposing plant or animal remains, manure, etc.).
Xy
xylophagous
Feeds on wood or bark; often species that develop in woody material.
Code
Definition
P
plant pest
The species damages living plants (agricultural crops, forest trees, ornamental plants).
S
stored-product pest
Attacks stored food or feed (grain, flour, seeds, etc.).
B
possibility of biodiversity influence
The species may influence native communities (competition, predation, habitat structure change, etc.).
Overview of alien species in the Czech Republic
Family / Taxon
Origin status
First occurrence
Taxon place of origin
Method of introduction
Invasive status
Food requirements
Note
Anthicidae
Omonadus floralis (Linnaeus 1758)
A
1951
S-AS
A
forest edges, hedgerows, dry pastures and scrubland
Stricticomus tobias (De Marseul 1879)
A
1944
AS
A
rotten plant tissue
Anthribidae
Araecerus fasciculatus (De Geer, 1775)
A
< 1900
AS
A
S
Ph(S)
corn, cassava, sweet potatoes, nutmeg, dried fruit, various nuts, coffee, tea
Gonotropis dorsalis (Gyllenhal, 1813)
A
Bostrichidae
Dinoderus minutus (Fabricius, 1775)
A
1965
?SE-AS
A
CU
Sa,Xy
imported bamboo, manioc (Cassava)
Lyctus brunneus (Stephens, 1830)
A
< 1850
SE-AS
A
CU
Xy
white wood, manioc
Rhyzopertha dominica (Fabricius, 1792)
A
1963
?SE-AS
A
S
Po(S)
cereals
Brentidae
Alocentron curvirostre (Gyllenhal 1833)
A
> 1904
AS
A
Ph(S)
Alcea rosae (Malvaceae)
Aspidapion (Aspidapion) validum (Germar 1817)
A
> 1960
AS
A
Ph(S)
Alcea rosae (Malvaceae)
Kalcapion semivittatum (Gyllenhal, 1833)
E
1999
MED
AS
IR
Ph
Mercurialis annua
Rhopalapion longirostre (Olivier 1807)
A
1999
SW-AS
AS
IR
Ph
Alcea rosae (Malvaceae)
Buprestidae
Lamprodila festiva festiva (Linnaeus, 1767)
E
2016
MED
A
juniper (Juniperus), cypress (Cupressus), thuja (Thuja), arborvitae (Chamaecyparis), platycladus (Platycladus)
Carabidae
Miscodera arctica (Paykull, 1798)
E
1984
N-EU
A
NA
Pr
Perigona nigriceps (Dejean, 1831)
A
1996
AS
A
compost, predator, gardens; also in peanuts
Pterostichus caspius (Ménétriés, 1832)
A
1980
SW-AS
A
NA
Pr
Cerambycidae
Monochamus sartor urussovii (Fischer von Waldheim, 1806)
A
1981
N-AS
A
C
Xy
Pinaceae
Morimus asper funereus (Mulsant, 1862)
E
1890
MED
A
C
Xy
deciduous trees
Nathrius brevipennis (Mulsant, 1839)
E
1908
MED
A
NH
Xy
Salix
Neoclytus acuminatus (Fabricius, 1775)
A
1950
C-AM
A
Ph
elm, ash, walnut (Ulmus, Fraxinus, Juglans)
Phymatodes lividus (Rossi, 1794)
E
1959
MED
A
(S)
Xy
oak, chestnut (Quercus, Castanea)
Semanotus russicus russicus (Fabricius, 1777)
E
2016
MED
A
juniper (Juniperus)
Trichoferus griseus (Fabricius, 1793)
E
1959
MED
A
C
Xy
fig tree, pistachio tree, rose (Ficus, Pistacia, Rosa)
Ciidae
Xylographus bostrichoides (Dufour, 1843)
A
1990
?AS
A
Fungi
Cleridae
Enoplium serraticorne (Olivier, 1790)
E
1990
W-MED
A
CR
Pr
Tarsostenus univittatus (Rossi, 1792)
C
1990
AUS
A
IA
Pr
Predator Bostrychidae and Anobiidae
Coccinellidae
Bulaea lichatschovii (Hummel, 1827)
A
?
AF/S-AS
A
sugar beet, lentils, alfalfa
Cryptolaemus montrouzieri (Mulsant, 1853)
A
1992
AUS
D
(S)
Pr
Coccinea
Delphastus catalinae (Horn, 1895)
A
1993
AM
D
(S)
Pr
Aleyrodidae
Harmonia axyridis (Pallas, 1773)
A
2006
E-AS
Pr
Aphids, Coccids
Hippodamia convergens (Guerin-Meneville, 1842)
A
1992
N-AM
D
(S)
Pr
Aphids
Cryptophagidae
Atomaria lewisi (Reitter, 1877)
A
1961
E-AS
S
IR
Sa
Caenoscelis subdeplanata (C.Brisout de Barneville, 1882)
A
1950
N-AM
mycophagous; forests; in decaying wood and plant material
Cryptophagus acutangulus (Gyllenhall, 1828)
C
1956
?
soil, mills
Cryptophagus cellaris (Scopoli, 1763)
C
1939
?
mycophagous, stored products, insect collections, herbariums
Cryptophagus fallax (Balfour-Browne, 1953)
C
< 1900
?
A
CU
Sa
Saved products
Cryptophagus laticollis (P. H. Lucas, 1846)
C
Cryptophagus simplex (Miller, 1858)
?
?
?
A
CU
Sa
moulds
Cryptophagus subfumatus (Kraatz, 1856)
C
1956
?
A
dried fruit, nuts
Curculionidae
Coccotrypes dactyliperda (Fabricius, 1801)
?
< 1900
?
A
(S)
Ph
Cyclorhipidion bodoanum (Reitter, 1913)
A
2020
AS
A
X
deciduous trees (mainly Quercus)
Dryocoetes himalayensis (Strohmeyer, 1908)
A
2009
S-AS
A
Xy
Juglans
Euophryum confine (Broun 1880)
A
1987
NZE
A
(S)
Xy
decaying wood
Gnathotrichus materiarius (Fitch 1858)
A
1933
N-AM
Xy
Picea, Pinus
Hylastinus fankhauseri (Reitter 1894)
E
1955
S-EU
A
C
Xy
Laburnum
Hypothenemus hampei (Ferrari 1867)
A
1988
C-AF
A
(S)
Ph
Coffea
Lignyodes bischoffi (Blatchley, 1916)
A
?
N-AM
A
Orthotomicus robustus (Knotek, 1899)
E
1979
MED
A
NH
Ph
Pinus nigra
Otiorhynchus armadillo (Rossi, 1792)
E
EU
S
Ph
Prunus, Ilex
Pachyrhinus lethierryi (Desbrochers des Loges, 1875)
E
?
MED
A
Chamaecyparis, Thuja, Cupressus (cypress, thuja, juniper)
Phloeotribus caucasicus (Reitter 1891)
A
1988
SW-AS
A
NH
Xy
Fraxinus
Pityogenes bistridentatus (Eichhoff, 1878)
E
< 1900
MED
A
NH
Xy
Pinus nigra
Sitophilus granarius (Linnaeus, 1758)
A
1350
SW-AS
A
S
Ph(S)
Seeds: wheat, barley, rye, oats, corn, rice, buckwheat, chestnuts, acorns
Sitophilus oryzae (Linnaeus 1763)
A
1896
S-AS
A
S
Ph(S)
Seeds: rice, wheat, barley, rye, oats, corn, rice, buckwheat, millet
Sitophilus zeamais (Motschulsky 1855)
C
1962
S-AS
A
S
Ph(S)
Seeds: rice, wheat, maize
Trypodendron laeve Eggers 1939
A
1987
AS
A
Xy
Coniferous trees
Xyleborinus attenuatus (Blandford, 1894)
A
1987
E-AS
A
NA
Xy
willow, oak, alder, birch, beech
Xyleborus pfeilii (Ratzeburg, 1837)
A
1837
AS
A
Xy
deciduous trees
Xyleborus volvulus (Fabricius, 1794)
A
1875
CN-AM
A
C
Xy
Xylosandrus germanus (Blandford, 1894)
A
2009
A-AS
A
Xy
deciduous and coniferous trees
Xylosandrus morigenus (Blandford 1894)
A
1916
AS
A
Xy
greenhouse orchids
Dermestidae
Anthrenocerus australis (Hope, 1843)
A
AUS
Anthrenus angustefasciatus (Ganglbauer, 1904)
E
2007
MED
Anthrenus flavipes LeConte, 1854
C
2001
N-AM
A
S
Sa
Anthrenus oceanicus Fauvel, 1903
A
2004
AUS
A
S
Sa
Attagenus doricus (Zhantiev, 2007)
A
2006
SE-EU
Attagenus lobatus (Rosenhauer, 1856)
A
2010
S-AS
A
damages textiles, carpets, clothing, museum collections
Attagenus smirnovi (Zhantiev, 1973)
C
1984
E-AF
A
(S)
Po
Attagenus unicolor (Brahm 1791)
C
Attagenus woodroffei (Halstead & Green, 1979)
E
2020
N-EU
A
pasta
Dermestes ater (De Geer, 1774)
A
1948
AM
A
CU
Sa
necrophagous
Dermestes haemorrhoidalis (Küster, 1852)
E
EU
A
S
Po(S)
Pests in warehouses and shops selling fur and leather products, cheese and dried meat
Dermestes peruvianus (Laporte de Castelnau, 1840)
A
1999
S-AM
A
(S)
Sa
animal products, textiles, corn; necrophagous
Megatoma ruflcornis (Aubé, 1866)
E
2016
EU
A
Orphinus fulvipes (Guérin-Méneville, 1838)
A
CN-AM
A
dried fruits, seeds, …
Reesa vespulae (Milliron, 1939)
A
1986
N-AM
A
S
Po(S)
Households and museum collections
Thylodrias contractus (Motschulsky, 1839)
E
2000
EU
A
S
Pa
Blattidae
Trogoderma angustum (Solier, 1849)
A
1991
S-AM
A
S
Household
Households and museum collections
Trogoderma glabrum (Herbst, 1783)
C
1904
A
households and solitary bee nests
Trogoderma granarium (Everts, 1898)
A
1962
S-AS
A
S
Po(S)
cereals and cereal products, groundnut seeds
Trogoderma inclusum (LeConte, 1854)
A
1956
N-AM
A
Sa(S)
necrophagous, stored products
Trogoderma longisetosum (Chao & Lee, 1966)
A
1967
AS
A
S
Po(S)
Trogoderma quinquefasciatum (Jacquelin, 1859)
A
< 1900
C-EU
A
S
Po
Trogoderma sternale sternale (Jayne, 1882)
A
?
?
Trogoderma variabile (Ballion, 1878)
A
1975
C-AS
A
S
Po(S)
wheat, dry plant products
Erotylidae
Dacne picta (Crotch, 1873)
A
1997
E-AS
A
C
Po(S)
fungi (Fungi)
Pharaxonotha kirschii (Reitter, 1875)
C
1900
C-AM
A
(S)
Sa
saprophage; grain, flour
Hydrophilidae
Cercyon inquinatus (Wollaston, 1854)
A
?
AF
A
Sa
carcasses, rotting deposits at the edges of water bodies, decaying leaves, rotting fruit, bat guano
Cercyon laminatus (Sharp, 1873)
A
1950
E-AS
A
IR
Sa
Cattle manure, leaf litter, compost, straw
Cercyon nigriceps (Marsham, 1802)
A
< 1930
ES-AS
A
Sa
herbivorous mammal droppings, rotting plant debris, compost, carrion, bird and small rodent nests
Cryptopleurum subtile (Sharp, 1884)
E
1952
E-AS
A
IR
Sa
decaying plant debris, compost, faeces of various mammals
Chrysomelidae
Acanthoscelides obtectus (Say, 1831)
A
< 1950
CS-AM
A
S
Ph(S)
legumes (beans, peas, chickpeas, lentils, soybeans, etc.)
Acanthoscelides pallidipennis (Motschulsky, 1874)
A
1992
N-AM
A
IH
Ph
Amorpha fructicosa
Bruchidius siliquastri (Delobel, 2007)
C
2003
?
A
I
Ph
Cercis siliquastrum
Bruchus ervi (Frölich, 1799)
E
1950
MED
A
S
Ph(S)
Lens
Bruchus lentis (Froelich, 1799)
A
I
Ph
Lens culinaris (lentil)
Bruchus pisorum (Linnaeus, 1758)
E
1850
MED
A
NH
Ph(PS)
Pisum sativum, Lathyrus sp., Vicia sp. (pea, vetch, vetchling)
Bruchus rufimanus (Bohemann, 1833)
A
1900
N-AF
I
Ph
Lathyrus venetus, Phaseolum vulgaris, Vicia, Cicer, Lens, Lupinus, Pisum
Bruchus signaticornis (Gyllenhal, 1833)
E
< 1900
MED
A
S
Ph(S)
Lens
Callosobruchus chinensis (Linnaeus, 1758)
A
1900
AS
A
S
Ph(S)
legumes (peas, chickpeas, beans, lentils, lotus, broad beans, etc.)
Callosobruchus maculatus (Fabricius, 1775)
A
1900
?
A
S
Ph
legumes (lentils, peas, broad beans, beans, etc.)
Callosobruchus phaseoli (Gyllenhal, 1833)
A
1945
AS
A
S
Ph
legumes
Caryedon gonagra (Fabricius, 1798)
A
?
?AF/SE-AS
A
(S)
Ph
Fruits of Indian tamarind (Tamarindus indica)
Caryedon serratus (Olivier, 1790)
A
1900
?AF/SE-AS
A
(S)
Ph(S)
oilseeds
Caryopemon cruciger (Stephens, 1839)
A
C
Ph
Abrus precatorius
Diabrotica virgifera virgifera (LeConte, 1868)
A
2002
N-AM
A
IH
Ph(P)
Zea (maize)
Leptinotarsa decemlineata (Say, 1824)
A
1945
N-AM
A
IH
Ph(P)
Solanum (potatoes), aubergine
Luperomorpha xanthodera (Fairmaire, 1888)
A
2012
AS
A
Po
herbs and ornamental plants (Lamiaceae, Brassicaceae, Asteraceae)
Megabruchidius dorsalis (Fahraeus, 1839)
A
1989
E-AS
A
S
Ph
Seeds of honey locust (Gleditsia)
Pseudopachymerina spinipes (Erichson, 1834)
A
?
S-AM
A
(S)
Ph
Acacia, Fabaceae
Zabrotes subfasciatus (Bohemann, 1833)
A
1987
C-AM
A
S
Ph(S)
legumes (beans, peas, etc.)
Kateretidae
Brachypterolus vestitus (Kiesenwetter, 1850)
E
1987
W-MED
S
IU
Ph
Anthirrhium
Laemophloeidae
Cryptolestes capensis (Waltl, 1834)
E
*1962
?S-AF
A
S
Po(S)
flour, spices, …
Cryptolestes duplicatus (Waltl 1839)
C
1990
?
A
under oak bark, stored products
Cryptolestes ferrugineus (Stephens, 1831)
C
< 1875
?
A
NU
Po(S)
cereals and products thereof
Cryptolestes pusilloides (Steel & Howe, 1952)
C
1978
?AUS
A
Po(S)
cereals, legumes, corn, spices, coffee, dried fruits, etc.
Cryptolestes pusillus (Schönherr, 1817)
A
< 1875
?
A
S
Po(S)
Flour, cereals and products thereof, coffee, cocoa beans, spices, dried fruit
Cryptolestes turcicus (Grouvelle, 1876)
C
1962
?
A
S
Po(S)
flour, cereals, legumes
Latridiidae
Adistemia watsoni (Wollaston, 1871)
C
1959
?
A
S
Sa
Tamarindus seeds, dry fruits, mushrooms, herbarium
Cartodere bifasciata (Reitter, 1877)
A
2000
A
A
Sa
barley malt, grass seeds, raisins, tobacco, wheat flour, etc.
Cartodere nodifer (Westwood, 1839)
A
1946
AUS
A
SC
Sa
mycophagous; compost, soil, hay
Corticaria elongata (Gyllenhal 1827)
C
1889
?
A
Sa
logs of deciduous trees, mouldy haystacks
Dienerella costulata (Reitter, 1877)
C
1900
?
A
S
Sa
food, roots, cellars, flats
Dienerella filum (Aubé, 1850)
C
1850
?
A
SC
Sa
cereals, herbariums, yeast, fungi, decomposing plant material
Lophocateridae
Ancyrona japonica (Reitter, 1889)
A
E-AS
A
Lophocateres pusillus (Klug, 1832)
A
1962
SE-AS
A
(S)
Sa
Saprophagous, necrophagous; rice, stored products
Mycetophagidae
Litargus balteatus (Leconte, 1856)
A
1983
AM
A
NA
Sa
compost, silage, rotting piles of chaff and grain, piles of fermenting fruit
Nitidulidae
Carpophilus dimidiatus (Fabricius, 1792)
A
1870
?S-AS
A
(S)
Sa(S)
dry fruits
Carpophilus hemipterus (Linnaeus, 1758)
A
1983
?S-AS
A
SIA
Sa(S)
dry fruits
Carpophilus ligneus (Murray, 1864)
A
1993
C-AM
A
(S)
Sa(S)
dry fruits
Carpophilus marginellus (Motschulsky, 1858)
A
1900
SE-AS
A
SIA
Sa(S)
nuts, cocoa beans, dill, dried coconut, pasta, spices, rice, peas, soybeans, corn, wheat, wheat flour, dried fruit
Carpophilus mutilatus (Erichson, 1843)
A
< 1900
?S-AS
A
(S)
Sa(S)
dry fruits
Carpophilus obsoletus (Erichson, 1843)
A
1985
SE-AS
A
(S)
Sa(S)
dry fruits, cereals
Carpophilus truncatus (Murray, 1864)
A
1983
SE-AS
A
SIR
Sa(S)
Glischrochilus quadrisignatus (Say, 1835)
A
1954
N-AM
A
IR
After
Fruit and vegetables
Nitidula flavomaculata (Rossi, 1790)
E
< 1900
MED
A
(S)
Sa
occasionally introduced with hides
Stelidota geminata (Say, 1825)
A
2012
N-AM
A
IR
After
fruit (mainly strawberries)
Ptiliidae
Acrotrichis insularis (Maklin, 1852)
A
2000
N-AM
A
IA
Sa
Coniferous and mixed forests (leaf litter, mosses, compost, decaying fungi)
Baeocrara japonica (Matthews, 1884)
A
1993
E-AS
A
Sa
manure, compost
Ptilodactylidae
Ptilodactyla exotica (Chapin, 1927)
C
1980
?
A
Ph
greenhouse-grown plants
Ptinidae
Epauloecus unicolor (Piller and Mitterpacher, 1783)
C
1900
barns, cattle stables, animal burrows
Gibbium psylloides (Czempinski, 1778)
C
1900
MED
A
S
Po(S)
Houses, hotels, stored products
Lasioderma serricorne (Fabricius, 1792)
A
1848
?
A
S
Sa(S)
Nicobium castaneum (Olivier, 1790)
C
1807
?
A
(S)
Sa
softwood, furniture, old books
Niptus hololeucus (Faldermann, 1836)
E
1500
E-MED
A
S
Sa
Ptinus bicinctus (Sturm 1837)
C
1856
?MED
A
warehouses, dwellings, old wooden objects, stored products
Ptinus exulans (Erichson, 1842)
A
1872
?AUS
A
CU
Sa
Ptinus fur (Linnaeus 1758)
C
1940
?
waste, dried vegetables
Ptinus latro (Fabricius, 1775)
C
1850
?
A
S
Sa
old wood
Ptinus tectus (Boieldieu 1856)
A
1937
AUS
A
S
Sa
Stored products
Tricorynus tabaci (Guérin-Méneville, 1850)
A
1965
C-AM
A
(S)
Ph
seeds
Trigonogenius globulus (Solier, 1849)
A
1939
CS-AM
A
S
Sa
dried animal products, insect collections, herbarium, stored products
Silvanidae
Ahasverus advena (Waltl, 1832)
C
1875
?N-AM
A
SC
Po(S)
saprophagous; stored products, compost
Oryzaephilus mercator (Fauvel, 1889)
A
1962
?
A
S
Po(S)
Oilseeds, tropical and subtropical nuts
Oryzaephilus surinamensis (Linnaeus, 1758)
C
1894
?
A
S
Po(S)
Rice, cereals, dry baked goods, breadcrumbs, oilseeds
Staphylinidae
Bisnius palmi (Smetana, 1955)
A
?
N-AM
Pr
Bohemiellina flavipennis (Cameron, 1920)
C
1941
?
A
Pr
grass compost
Lithocharis nigriceps (Kraatz, 1859)
A
1912
SE-AS
A
IA
Pr
compost, predator
Nacaeus impressicollis (Motschulsky, 1857)
A
2005
AF
A
Pr
Oxytelus migrator (Fauvel, 1904)
A
1978
SE-AS
A
IA
Pr
hay, grass, compost, manure, horse excrement
Philonthus ohizumi (R. Dvořák, 1958)
A
2012
E-AS
A
IA
Pr
animal excrement, carcasses
Philonthus rectangulus (Sharp, 1874)
A
1933
E-AS
A
IA
Pr
compost, predator
Philonthus spinipes spinipes Sharp, 1874
A
1982
E-AS
A
IR
Pr
animal excrement, compost, manure, carrion, decaying organic plant debris
Trichiusa immigrata (Lohse, 1984)
A
1989
N-AM
A
IR
Pr
compost, predator, fungi
Tenebrionidae
Alphitophagus bifasciatus (Say, 1824)
C
1940
?
A
CA
Sa
remnants; compost, rotten fruit; under the bark of old stumps
Blaps gigas (Linnaeus, 1758)
E
1888
MED
A
CU
Sa
Gnatocerus cornutus (Fabricius, 1798)
A
1900
?S-AM
A
S
Sa(S)
cereals in storage
Latheticus oryzae (Waterhouse, 1880)
A
1973
?S-AS
A
(S)
Sa
stored products, cereals in warehouses
Lyphia tetraphylla (Fairmaire, 1856)
A
1934
N-AM
A
CA
Sa
Palorus subdepressus (Wollaston, 1864)
A
1983
?S-AM
A
NA
Sa
stored products, cereals, tree fungi
Tribolium castaneum (Herbst, 1797)
C
1900
?
A
S
Sa(S)
Stored products
Tribolium confusum (Jacquelin du Val, 1861)
A
1900
AF
A
S
Sa(S)
stored products
Tribolium destructor (Uyttenboogaart, 1933)
A
1948
S-AS
A
S
Sa(S)
mills, agricultural product warehouses, households
Thanerocleridae
Thaneroclerus buqueti (Lefebvre, 1835)
A
1963
S-AS
A
(S)
Pr
Trogossitidae
Tenebroides mauritanicus (Linnaeus, 1758)
A
1803
AF
A
*NA
Sa
Saprophagous, carnivorous; stored products
Zopheridae
Pycnomerus inexpectus (Jaquelin Du Val, 1859)
C
1901
?
A
orchids (greenhouses)