Carabidae

Details: Published: 19 January 2026

Bioindication is a key tool in modern ecology, allowing us to read the state of the environment through living organisms. While physical and chemical measurements provide us with an instant "snapshot" of air or water quality, bioindicators – organisms that are sensitive to environmental changes – give us information about the long-term state and stability of the entire ecosystem. Among terrestrial invertebrates, the family Carabidae occupies an important position in this regard.

Ground beetles are abundant, species-rich and well researched. They often respond to moisture and temperature gradients, soil surface structure and the degree of habitat degradation. Monitoring is mainly carried out using ground traps, which allow standardised data collection across time and space. The bioindication system for Carabidae in the Czech Republic has gradually developed on the basis of long-term research. Currently, species are categorised according to their habitat dependence and sensitivity to environmental changes as relict (R), adaptable (A) and eurytopic (E). These categories facilitate the interpretation of field data and enable effective ecological assessment of sites.

The quality of a biotope is not assessed solely on the basis of the presence of a single "rare" beetle, but on the structure of the entire community. The following are usually monitored:

Ratio of ecological groups: The predominance of R and E species indicates a well-preserved habitat.
Diversity index: A higher number of species with even representation indicates a balanced ecosystem.
Presence of specialists: For example, the occurrence of wingless (brachypterous) species indicates long-term habitat continuity, as these beetles are unable to move quickly to another location if the site is destroyed.

The following paragraphs explain the individual columns in the table below.

Taxon

Bioindication group

Habitat type

The biotope types were taken from lists originating from this source: P. Vonička et al.: Lists of indicator species of animals and fungi for individual types of natural habitats according to the Czech Republic biotope catalogue, Ekologické služby, September 2016

The following table provides an overview of the individual habitats used:

Code	Habitat name
1340	Inland salt meadows
2330	Open grasslands of continental dunes with Corynephorus and Agrostis
3130	Oligotrophic to mesotrophic standing waters with vegetation of the Littorelletea uniflorae or Isoëto-Nanojuncetea classes
3220	Alpine rivers and herbaceous vegetation along their banks
3230	Alpine rivers and their woody vegetation with German broom
3240	Alpine rivers and their woody vegetation with grey willow
3270	Muddy river banks with vegetation of the Chenopodion rubri p. p. and Bidention p. p. associations.
4030	European dry heaths
4060	Alpine and boreal heaths
4070	Scrub with mountain pine and rhododendron
4080	Subarctic willow scrub
40A0	Continental deciduous scrub
5130	Juniper formations on heaths or calcareous grasslands
6110	Calcareous or alkaline rocky grasslands
6150	Silicate alpine and boreal grasslands
6190	Pannonian rocky grasslands
6210	Semi-natural dry grasslands and scrubland facies on calcareous substrates
6230	Species-rich siliceous grasslands on siliceous substrates in mountain and sub-mountain areas
6240	Sub-Pannonian steppe grasslands
6250	Pannonian loess steppe grasslands
6260	Pannonian sand steppes
6410	Tall grasslands on calcareous, peaty or clayey soils
6440	Floodplain meadows of river valleys of the Cnidion dubii association
6510	Lowland hay meadows
6520	Mountain hay meadows
7110	Active raised bogs
7120	Degraded raised bogs (capable of natural regeneration)
7140	Transitional mires and quaking bogs
7150	Depressions on peat substrate
7210	Calcareous fens with Cladium mariscus and species of the Caricion davallianae association
7230	Alkaline fens
8110	Silicate scree of mountain to nival level
8150	Central European silicate scree
8160	Calcareous scree of the hilly and mountainous levels
8210	Calcareous rocky slopes with chasmophytic vegetation
8220	Silicate rocky slopes with chasmophytic vegetation
8230	Pioneer vegetation on silicate rocks
9110	Beech forests of the Luzulo-Fagetum association
9130	Beech forests of the Asperulo-Fagetum association
9140	Central European subalpine beech forests with maple trees
9150	Central European limestone beech forests
9170	Oak-hornbeam forests of the Galio-Carpinetum association
9180	Tilio-Acerion forests on slopes, scree and ravines
9190	Old acidophilous oak forests with pedunculate oak on sandy plains
91D0	Peat bog woodland
91E0	Mixed ash-alder floodplain forests of temperate and boreal Europe
91F0	Mixed floodplain forests with English oak, European white elm, European white elm, European ash or narrow-leaved ash along major rivers
91G0	Pannonian oak-hornbeam forests
91H0	Pannonian oak-hornbeam forests
91I0	Euro-Siberian steppe oak forests
91T0	Central European lichen pine forests
91U0	Forest-steppe pine forests
9410	Acidophilic spruce forests of the mountain to alpine level
N/A	Sandy, sandy loam and clayey river banks without vegetation or with sparse vegetation (an important habitat for a number of endangered species closely linked to this disappearing habitat, which is not listed in the types of natural habitats)

Ecological group

Ecological groups were taken from lists originating from this source: P. Vonička et al.: Lists of indicator species of animals and fungi for individual types of natural habitats according to the Czech Republic biotope catalogue, Ecological Services, September 2016

The terms listed are used in entomology (and ecology in general) to describe the ecological relationship of insects to a particular environment or specific conditions (soil, humidity, salinity).

The suffixes are of fundamental importance:

-biont: The organism is strictly bound to a given environment and cannot exist without it.
-phile: The organism prefers a given environment but is able to survive elsewhere.

The following ecological groups are used in the table of bioindicators:

Term	Description
Link to saline environments (salt marshes) This group includes species living in places with high salt content.
Halobiont	A species living exclusively in saline habitats (e.g. inland salt marshes, sea coasts). It dies without salt.
Halophile	A species that seeks out salty environments but also occurs in habitats with low salinity.
Subhalophile	A species that tolerates salt and often occurs on the edges of salt marshes, but is not directly dependent on it.
Connection to moisture and water
Hygrophile	A moisture-loving species. Requires a high level of humidity (e.g. floodplain forests, marshes).
Ripicole	A species bound to the banks of water bodies (rivers, streams, lakes). Often lives in sediment or under stones near water.
Subripicole	A species that prefers proximity to water, but can also be found in the wider vicinity of riparian vegetation, not directly at the water's edge.
Substrate dependence (sand)
Psamobiont	A species exclusively associated with sandy substrates (e.g. dunes, drifting sands). It cannot complete its development in other types of soil.
Psamophile	A species that prefers sand (e.g. because it is easy to burrow in), but is also able to live in other light soils.
Bound to peat bogs
Tyrphobiont	Species critically bound to peat bogs. These are often glacial relicts (remnants from the Ice Age) that cannot survive elsewhere.
Tyrphophile	A species that finds optimal conditions in peat bogs, but can also inhabit other acidic and waterlogged areas.
Connection to heathland
Callunobiont	A species that is dependent on heathland for food or habitat (often monophagous insects that feed on heather).
Callunophile	A species that prefers heathlands, but also occurs in dry coniferous forests or on dry slopes with similar characteristics.

Status in habitat

Occurrence in the Czech Republic

Sources

Bioindicator groups of the families Carabidae and Cicindelidae in the Czech and Slovak Republics
Taxon	Bioindication group	Habitat Type	Ekological group	Status in habitat	CZ
Abax carinatus carinatus	A	91		Konst	•
Abax elatus elatus	A	40A0		Konst	•
Abax ovalis	A	9110, 9130, 9140, 9150, 9180		Konst	•
Abax parallelepipedus parallelepipedus	A	91F0		Konst	•
Abax parallelus parallelus	A	9170, 9190, 91G0, 91H0, 91I0		Konst	•
Abax piceus piceus	A	9110, 9130, 9140, 9150, 9180		Konst	•
Abax schueppeli rendschmidtii	R	9110, 9130, 9140, 9150, 9180		Dg	•
Acinopus picipes	A
Acupalpus brunnipes	R	3130	hygrophile	Dg, Rar	•
Acupalpus dubius	R	3130	hygrophile	Dg, Rar	•
Acupalpus elegans	R	1340	halobiont	Dg, Rar	•
Acupalpus exiguus	A	3130	hygrophile	Konst	•
Acupalpus flavicollis	A				•
Acupalpus interstitialis	R	6210		Dg	•
Acupalpus luteatus	A	3130	hygrophile	Konst, Rar	•
Acupalpus maculatus	A	1340	halophile	Konst	•
Acupalpus meridianus	E				•
Acupalpus parvulus	A	3130	hygrophile	Konst	•
Acupalpus suturalis	R	1340	halophile	Dg, Rar	•
Agonum angustatum	A	3130	hygrophile	Dg
Agonum antennarium	R				•
Agonum dolens	R	3130	hygrophile	Dg, Rar	•
Agonum duftschmidi	A	3130	hygrophile	Konst	•
Agonum emarginatum	A	3130	hygrophile	Konst	•
Agonum ericeti	R	7110, 7120, 7140, 91D0	tyrphobiont	Dg	•
Agonum fuliginosus	A				•
Agonum gracile	A	7110, 7120, 7140, 7150, 91D0	hygrophile	Konst	•
Agonum gracilipes	E				•
Agonum hypocrita	R	7210, 7230	hygrophile	Dg, Rar	•
Agonum impressum	R
Agonum longicorne	R
Agonum lugens	R	3130, 7210, 7230	hygrophile	Dg	•
Agonum marginatum	A				•
Agonum micans	A	3270, 91E0	ripicole, hygrophile	Konst	•
Agonum monachum monachum	R	1340	halophile	Dg, Rar	•
Agonum muelleri	E				•
Agonum permoestum	R
Agonum piceum	A	3130	hygrophile	Konst	•
Agonum sexpunctatum	A				•
Agonum thoreyi	A				•
Agonum versutum	A	3130	hygrophile	Konst	•
Agonum viduum	A				•
Agonum viridicupreum viridicupreum	A	3130	hygrophile	Konst, Rar	•
Amara aenea	E				•
Amara anthobia	R	40A0, 6210		Dg, Rar	•
Amara apricaria	E				•
Amara aulica	E				•
Amara bifrons	E				•
Amara brunnea	A	91D0	hygrophile	Konst	•
Amara communis	A	91D0		Konst	•
Amara concinna	A	6410, 6440, 6510		Konst, Rar	•
Amara consularis	E				•
Amara convexior	E				•
Amara convexiuscula	E				•
Amara crenata	R	6210, 6240, 6250		Dg, Rar	•
Amara cursitans	E				•
Amara curta	A	40A0, 5130, 6110, 6190, 6230, 6520, 91T0, 91U0		Konst	•
Amara equestris equestris	A	5130, 6110, 6190		Konst	•
Amara erratica	R	4060, 6150, 6230, 6520		Dg	•
Amara eurynota	E				•
Amara famelica	A	4030, 5130, 91T0, 91U0		Konst	•
Amara familiaris	E				•
Amara fulva	A	2330	psamophile	Konst	•
Amara fulvipes	R				•
Amara fusca	R				•
Amara gebleri	A	6410, 6440, 6510		Konst	•
Amara chaudoiri incognita	A	6410, 6440, 6510		Konst, Rar	•
Amara infima	R	4030, 5130	callunophile	Dg	•
Amara ingenua	E				•
Amara littorea	E				•
Amara lucida	A	6210, 6240, 6250		Konst	•
Amara lunicollis	A	6230, 6520		Konst	•
Amara majuscula	E				•
Amara makolskii	A	91D0		Konst	•
Amara montivaga	E				•
Amara municipalis municipalis	E				•
Amara nigricornis	R	9410		Dg, Rar	•
Amara nitida	A				•
Amara ovata	E				•
Amara plebeja	E				•
Amara praetermissa	A	4030, 4060, 5130, 6150, 6230, 6520		Konst	•
Amara proxima	R
Amara pulpani	R	4030, 4060, 5130, 6110, 6190		Dg	•
Amara quenseli quenseli	R
Amara sabulosa	E				•
Amara saginata	R
Amara saphyrea	A
Amara schimperi	R	3220/30/40	ripicole	Dg, Rar	•
Amara similata	E				•
Amara sollicita	R
Amara spreta	A	2330, 6260	psamobiont	Konst	•
Amara strandi	R	1340	halobiont	Dg, Rar	•
Amara strenua	R				•
Amara tibialis	A				•
Amara tricuspidata	A	6260		Konst	•
Amblystomus niger	R	1340, 6410, 6440, 6510	subhalophile	Dg, Rar	•
Anchomenus dorsalis	E				•
Anisodactylus binotatus	E				•
Anisodactylus nemorivagus	A	5130		Konst	•
Anisodactylus poeciloides poeciloides	R				•
Anisodactylus signatus	E				•
Anthracus consputus	A	3130	hygrophile	Konst	•
Anthracus longicornis	R	1340	halophile	Dg	•
Aptinus bombarda	R	9110, 9130, 9140, 9150, 9180, 9170, 9190, 91G0, 91H0, 91I0		Dg	•
Asaphidion austriacum	A		ripicole	Konst
Asaphidion caraboides caraboides	R				•
Asaphidion flavipes	E				•
Asaphidion pallipes	A				•
Atranus ruficollis	R
Badister bullatus	A				•
Badister collaris	A	3130	hygrophile	Konst	•
Badister dilatatus	A	3130	hygrophile	Konst	•
Badister dorsiger	R	3130, 91E0	hygrophile	Dg, Rar	•
Badister lacertosus	A	91	hygrophile	Konst	•
Badister meridionalis	A	3130	hygrophile	Konst	•
Badister peltatus peltatus	A	3130, 7210, 7230	hygrophile	Konst, Rar	•
Badister sodalis	A	6410, 6440, 6510		Konst	•
Badister unipustulatus	A	3130	hygrophile	Konst	•
Bembidion argenteolum	R		ripicole	Dg, Rar	•
Bembidion articulatum	E				•
Bembidion ascendens	R	3220/30/40	ripicole	Dg, Rar	•
Bembidion assimile	A	3130	hygrophile	Konst	•
Bembidion atrocaeruleum	R	3220/30/40	ripicole	Dg	•
Bembidion azurescens azurescens	A	3220/30/40	ripicole	Konst	•
Bembidion biguttatum	A				•
Bembidion bipunctatum bipunctatum	R	3220/30/40, 6150	ripicole	Dg, Rar	•
Bembidion bruxellense	A				•
Bembidion bualei bualei	R	3220/30/40	ripicole	Dg, Rar	•
Bembidion conforme	R	3220/30/40	ripicole	Dg, Rar	•
Bembidion dalmatinum dalmatinum	R	9170, 9190, 91G0, 91H0, 91I0		Dg, Rar	•
Bembidion decorum decorum	A	3220/30/40	ripicole	Konst	•
Bembidion deletum	A				•
Bembidion dentellum	A	91	hygrophile	Konst	•
Bembidion doris	A	3130	hygrophile	Konst	•
Bembidion ephippium	R
Bembidion fasciolatum	R				•
Bembidion femoratum femoratum	E				•
Bembidion fluviatile fluviatile	A		ripicole	Konst	•
Bembidion foraminosum	R		ripicole	Dg, Rar	•
Bembidion friebi	R
Bembidion fulvipes	R				•
Bembidion fumigatum	A	1340, 3130, 7210, 7230	subhalophile	Konst	•
Bembidion genei illigeri	E				•
Bembidion geniculatum geniculatum	A	3220/30/40	ripicole	Konst	•
Bembidion gilvipes	A	6410, 6440, 6510		Konst	•
Bembidion glaciale	R
Bembidion guttula guttula	A	3130	hygrophile	Konst	•
Bembidion humerale	R	7110, 7120, 7140, 7150, 91D0	tyrphophile, hygrophile	Dg	•
Bembidion inoptatum	A	3130	hygrophile	Konst	•
Bembidion lampros	E				•
Bembidion laticolle	R		ripicole	Dg, Rar	•
Bembidion latiplaga	A
Bembidion litorale	R		ripicole	Dg	•
Bembidion lunatum	A	3220/30/40	subripicole	Konst, Rar	•
Bembidion lunulatum	A				•
Bembidion mannerheimi	A				•
Bembidion milleri carpathicum	A				•
Bembidion milleri milleri	A				•
Bembidion minimum	A	1340, 3130	subhalophile	Konst	•
Bembidion modestum	A	3220/30/40	ripicole	Konst, Rar	•
Bembidion monticola monticola	A	3220/30/40	ripicole	Konst	•
Bembidion neresheimeri neresheimeri	R	3130	subhalophile	Dg, Rar	•
Bembidion nigricorne	R	4030, 91T0, 91U0		Dg, Rar	•
Bembidion normannum normannum	R
Bembidion obliquum	A				•
Bembidion obtusum	E				•
Bembidion octomaculatum	A	3130	hygrophile	Konst	•
Bembidion prasinum	R	3220/30/40	ripicole	Dg, Rar	•
Bembidion properans	E				•
Bembidion punctulatum punctulatum	A	3220/30/40	ripicole	Konst	•
Bembidion pygmaeum	R		ripicole	Dg	•
Bembidion quadrimaculatum quadrimaculatum	E				•
Bembidion quadripustulatum quadripustulatum	A	3130	hygrophile	Konst	•
Bembidion ruficolle	R		ripicole	Dg, Rar	•
Bembidion scapulare lomnickii	R				•
Bembidion semipunctatum	A	3270	ripicole	Konst	•
Bembidion schueppelii	A	91	hygrophile	Konst	•
Bembidion splendidum	R				•
Bembidion stephensi	A				•
Bembidion striatum	R		ripicole	Dg, Rar	•
Bembidion subcostatum vau	R	3220/30/40	ripicole	Dg, Rar	•
Bembidion subfasciatum	R				•
Bembidion tenellum tenellum	A	3130	hygrophile	Konst	•
Bembidion testaceum testaceum	R	3220/30/40	ripicole	Dg, Rar	•
Bembidion tetracolum tetracolum	E				•
Bembidion tibiale	A	3220/30/40	ripicole	Konst	•
Bembidion varicolor varicolor	R	3220/30/40	ripicole	Dg, Rar	•
Bembidion varium	E				•
Bembidion velox	R		ripicole	Dg, Rar	•
Blemus discus discus	A				•
Blethisa multipunctata multipunctata	A	3130	hygrophile	Konst	•
Bradycellus caucasicus	A	4030, 4060, 5130		Konst	•
Bradycellus csikii	E				•
Bradycellus harpalinus	A	4070/80, 6150, 6230, 6520, 91D0		Konst	•
Bradycellus ruficollis	R	4030, 4060, 5130	callunobiont	Dg	•
Bradycellus verbasci	A				•
Brachinus bipustulatus	A
Brachinus crepitans	E				•
Brachinus elegans	A	6410, 6440, 6510		Konst, Rar	•
Brachinus explodens	E				•
Brachinus incertus	A
Brachinus plagiatus	A
Brachinus psophia	A	6410, 6440, 6510		Konst, Rar	•
Broscus cephalotes	A	2330	psamophile	Konst	•
Calathus ambiguus ambiguus	A				•
Calathus cinctus	A	6240, 6250, 6260		Konst	•
Calathus erratus erratus	A	4030, 91T0, 91U0		Konst	•
Calathus fuscipes fuscipes	E				•
Calathus melanocephalus melanocephalus	E				•
Calathus metallicus metallicus	A
Calathus micropterus	A	4030, 4060, 4070/80, 5130, 6150, 6230, 6520, 91D0		Konst	•
Callistus lunatus lunatus	A	6240, 6250		Konst	•
Calodromius spilotus	A
Calosoma inquisitor inquisitor	A	9170, 9190, 91G0, 91H0, 91I0		Konst	•
Calosoma maderae maderae	A				•
Calosoma reticulatum	R
Calosoma sycophanta	A	9170, 9190, 91G0, 91H0, 91I0		Konst	•
Carabus arvensis arvensis	A	4030, 4060, 5130, 6150, 6230, 6520, 7110, 7120, 7140, 91D0, 91T0, 91U0		Konst	•
Carabus arvensis eremita	A
Carabus auratus	A	6410, 6440, 6510		Konst	•
Carabus auronitens auronitens	A	9410		Konst	•
Carabus auronitens escheri	A
Carabus cancellatus cancellatus	A				•
Carabus cancellatus tibiscinus	A
Carabus clathratus auraniensis	R	91	hygrophile	Dg, Rar	•
Carabus convexus convexus	A				•
Carabus coriaceus coriaceus	A				•
Carabus coriaceus rugifer	A
Carabus fabricii fabricii	R				•
Carabus glabratus glabratus	A	9110, 9130, 9140, 9150, 9180, 9410		Konst	•
Carabus granulatus granulatus	E				•
Carabus hortensis hortensis	A				•
Carabus hungaricus hungaricus	R	6210, 6240, 6250		Dg, Konst, Rar	•
Carabus intricatus intricatus	A				•
Carabus irregularis irregularis	R	9110, 9130, 9140, 9150, 9180		Dg	•
Carabus irregularis montadoni	R	9110, 9130, 9140, 9150, 9180		Dg	•
Carabus linnei linnei	A	9410		Konst	•
Carabus menetriesi menetriesi	R
Carabus menetriesi pacholei	R	7110, 7120, 7140	tyrphobiont	Dg, Rar	•
Carabus montivagus montivagus	R
Carabus nemoralis nemoralis	A				•
Carabus nitens	R	4030, 7110, 7120, 7140		Dg, Rar	•
Carabus obsoletus obsoletus	A	9110, 9130, 9140, 9150, 9180		Konst, Rar	•
Carabus problematicus	A	4030, 91T0, 91U0		Konst	•
Carabus scabriusculus scabriusculus	A	6210, 6240, 6250		Konst, Rar	•
Carabus scheidleri helleri	A	6410, 6440, 6510		Konst	•
Carabus scheidleri scheidleri	A	6410, 6440, 6510		Konst	•
Carabus scheidleri zawadzkii	A
Carabus sylvestris sylvestris	A	4070/80, 6150, 91D0, 9410		Konst	•
Carabus sylvestris transsylvanicus	A
Carabus ulrichii rhilensis	A	9170, 9190, 91G0, 91H0, 91I0, 91F0		Konst, Rar	•
Carabus ulrichii ulrichii	A	6410, 6440, 6510, 9170, 9190, 91G0, 91H0, 91I0, 91F0		Konst	•
Carabus variolosus	R	9110, 9130, 9140, 9150, 9180	hygrophile	Dg, Rar	•
Carabus violaceus andrzejuscii	A
Carabus violaceus exasperatus	A				•
Carabus violaceus violaceus	A				•
Cephalophonus cephalotes	R				•
Cicindela campestris campestris	A				•
Cicindela hybrida hybrida	A	2330	psamophile	Konst	•
Cicindela littoralis nemoralis	R				•
Cicindela soluta pannonica	R	2330, 6260	psamophile	Dg, Rar	•
Cicindela sylvatica sylvatica	A	91T0, 91U0	psamobiont	Konst	•
Cicindela sylvicola	A				•
Clivina collaris	E				•
Clivina fossor	E				•
Clivina ypsilon	R				•
Cychrus attenuatus	R	9110, 9130, 9140, 9150, 9180		Dg	•
Cychrus caraboides caraboides	A				•
Cylindera arenaria viennensis	R	2330	psamophile	Dg	•
Cylindera germanica germanica	A	6240, 6250		Konst, Rar	•
Cymindis angularis	A	40A0, 6210, 6240, 6250, 6260		Konst	•
Cymindis axillaris axillaris	R	40A0, 6110, 6190		Dg, Rar	•
Cymindis cingulata	R	5130, 6230, 6520		Dg, Rar	•
Cymindis humeralis	A	5130, 6110, 6190, 6210		Konst	•
Cymindis macularis	R	2330, 6260	psamobiont	Dg, Rar	•
Cymindis miliaris	R	6110, 6190		Dg, Rar	•
Cymindis scapularis	R				•
Cymindis vaporariorum	R	4030, 4060, 7110, 7120, 7140, 7150, 91D0		Dg, Rar	•
Deltomerus carpathicus	R
Deltomerus tatricus	R
Demetrias atricapillus	E				•
Demetrias imperialis	A	3130	hygrophile	Konst	•
Demetrias monostigma	A				•
Diachromus germanus	A	3130, 6410, 6440, 6510	hygrophile	Konst	•
Dicheirotrichus cognatus	R	4030		Dg, Rar	•
Dicheirotrichus placidus	A	3130	hygrophile	Konst	•
Dicheirotrichus rufithorax	E				•
Dinodes decipiens decipiens	R
Dixus clypeatus	A
Dolichus halensis	E				•
Dromius agilis	A				•
Dromius angustus	A				•
Dromius fenestratus	A	9410		Konst	•
Dromius kuntzei	A				•
Dromius laeviceps	A	91F0		Konst, Rar	•
Dromius meridionalis	A
Dromius quadraticollis	A	91F0		Konst	•
Dromius quadrimaculatus	A				•
Dromius schneideri	A	91T0, 91U0		Konst	•
Drypta dentata	E				•
Duvaliopsis pilosella poloninensis	R
Duvaliopsis pilosella stobieckii	R	9410		Dg, Rar	•
Duvalius bokori bokori	R
Duvalius bokori broziki	R
Duvalius bokori gelidus	R
Duvalius bokori valyianus	R
Duvalius goemoeriensis	R
Duvalius hungaricus brzotinensis	R
Duvalius hungaricus hungaricus	R
Duvalius hungaricus slovacus	R
Duvalius hungaricus szilicezensis	R
Duvalius microphthalmus microphthalmus	R
Duvalius microphthalmus spelaeus	R
Duvalius microphthalmus tatricus	R
Duvalius microphthalmus vorago	R
Duvalius subterraneus subterraneus	R
Duvalius szaboi szaboi	R
Duvalius szaboi veporensis	R
Dyschirius abditus	R	3220/30/40	ripicole	Dg, Rar	•
Dyschirius aeneus aeneus	E				•
Dyschirius agnatus	A	3270	subripicole	Konst	•
Dyschirius angustatus	R				•
Dyschirius bonellii	R	6210, 6240, 6250		Dg, Rar	•
Dyschirius digitatus	A	3220/30/40	ripicole	Konst, Rar	•
Dyschirius extensus	R
Dyschirius globosus	E				•
Dyschirius gracilis	R
Dyschirius chalceus	R	1340	halobiont	Dg, Rar	•
Dyschirius chalybeus gibbifrons	A	1340	halophile	Konst, Rar	•
Dyschirius intermedius	A	3270	ripicole	Konst	•
Dyschirius laeviusculus	R	1340	subhalophile	Dg, Rar	•
Dyschirius latipennis	A				•
Dyschirius nitidus nitidus	A	3270	subripicole	Konst, Rar	•
Dyschirius parallelus ruficornis	A
Dyschirius politus politus	A	3270	subripicole	Konst	•
Dyschirius pusillus	R
Dyschirius rufipes	R	6210, 6240, 6250		Dg, Rar	•
Dyschirius salinus striatopunctatus	R				•
Dyschirius strumosus	R	1340	halobiont	Dg, Rar	•
Dyschirius substriatus priscus	R
Dyschirius thoracicus	A				•
Dyschirius tristis	A				•
Elaphrus aureus aureus	R		ripicole	Dg	•
Elaphrus cupreus	A				•
Elaphrus riparius	E				•
Elaphrus uliginosus	A	3130	hygrophile	Konst, Rar	•
Elaphrus ullrichi	R				•
Harpalus affinis	E				•
Harpalus akinini	A
Harpalus albanicus	R	6210, 6240, 6250		Dg	•
Harpalus anxius	A	2330	psamophile	Konst	•
Harpalus atratus	A	40A0		Konst	•
Harpalus autumnalis	A	2330, 6260, 91T0, 91U0	psamobiont	Konst	•
Harpalus calceatus	A				•
Harpalus caspius	A	6210		Konst	•
Harpalus cephalotes	R	1340	halophile	Dg, Rar	•
Harpalus cisteloides hurkai	R	6210		Dg	•
Harpalus cupreus fastuosus	R	3130	hygrophile	Dg, Rar	•
Harpalus dimidiatus	A
Harpalus distinguendus distinguendus	E				•
Harpalus flavescens	R	2330	psamobiont	Dg, Rar	•
Harpalus flavicornis flavicornis	R	6210, 6240, 6250		Dg, Rar	•
Harpalus froelichii	A	2330	psamophile	Konst	•
Harpalus fuscipalpis	R				•
Harpalus griseus	E				•
Harpalus hirtipes	R	2330, 6260	psamobiont	Dg, Rar	•
Harpalus honestus	A	6110, 6190		Konst	•
Harpalus hospes hospes	R	6210, 6240, 6250		Dg, Rar	•
Harpalus laevipes	A				•
Harpalus latus	A				•
Harpalus luteicornis	A	6410, 6440, 6510		Konst	•
Harpalus marginellus	A	9110, 9130, 9140, 9150, 9180, 9170, 9190, 91G0, 91H0, 91I0		Konst, Rar	•
Harpalus melancholicus	R				•
Harpalus melancholicus melancholicus	R	2330, 6260	psamobiont	Dg, Rar	•
Harpalus modestus	R	2330, 6260	psamophile	Dg, Rar	•
Harpalus neglectus neglectus	R	2330	psamobiont	Dg, Rar	•
Harpalus oblitus oblitus	A
Harpalus picipennis	R	2330, 6260	psamobiont	Dg	•
Harpalus politus politus	R	6210		Dg	•
Harpalus progrediens	R	91	hygrophile	Dg	•
Harpalus pumilus	A	6110, 6190		Konst	•
Harpalus pygmaeus	R				•
Harpalus rubripes	E				•
Harpalus rufipalpis	A	40A0, 91T0, 91U0		Konst	•
Harpalus rufipes	E				•
Harpalus saxicola	R
Harpalus serripes serripes	A				•
Harpalus servus	R	2330, 6260	psamobiont	Dg, Rar	•
Harpalus signaticornis	E				•
Harpalus smaragdinus	A				•
Harpalus solitaris	A	4030, 5130, 91D0, 9410		Konst	•
Harpalus subcylindricus	A				•
Harpalus tardus	E				•
Harpalus tenebrosus	A				•
Harpalus xanthopus winkleri	A	9170, 9190, 91G0, 91H0, 91I0		Konst, Rar	•
Harpalus zabroides	R	6210, 6240, 6250		Dg, Rar	•
Chlaenius festivus festivus	A				•
Chlaenius nigricornis	A	3130	hygrophile	Konst	•
Chlaenius nitidulus	A	3130	hygrophile	Konst	•
Chlaenius spoliatus	A	3270	ripicole	Konst	•
Chlaenius sulcicollis	R
Chlaenius tibialis	R
Chlaenius tristis tristis	A	3130	hygrophile	Konst	•
Chlaenius vestitus	A				•
Laemostenus terricola punctatus	A
Laemostenus terricola terricola	A				•
Lebia cruxminor cruxminor	A	40A0, 6210		Konst	•
Lebia cyanocephala cyanocephala	R	6210		Dg	•
Lebia humeralis	R	9170, 9190, 91G0, 91H0, 91I0		Dg, Rar	•
Lebia chlorocephala	A				•
Lebia marginata	R	9170, 9190, 91G0, 91H0, 91I0		Dg, Rar	•
Lebia scapularis scapularis	R
Lebia trimaculata	R
Leistus ferrugineus	E				•
Leistus montanus corconticus	R	8110, 8150, 8160, 8210, 8220, 8230		Dg, Rar	•
Leistus montanus kultianus	R	8110, 8150, 8160, 8210, 8220, 8230		Dg	•
Leistus montanus montanus	R				•
Leistus montanus pawlowskii	R
Leistus piceus piceus	A	6150, 9110, 9130, 9140, 9150, 9180, 9410		Konst	•
Leistus rousi	R
Leistus rufomarginatus	R	9170, 9190, 91G0, 91H0, 91I0, 91E0, 91F0		Dg	•
Leistus terminatus	A	3130	hygrophile	Konst	•
Licinus cassideus cassideus	R	6210, 6240, 6250		Dg, Rar	•
Licinus depressus	A				•
Licinus hoffmannseggii	R	9110, 9130, 9140, 9150, 9180		Dg	•
Limodromus assimilis	A				•
Limodromus krynickii	R	91	hygrophile	Dg, Rar	•
Limodromus longiventris	R	3130, 91E0	hygrophile	Dg, Rar	•
Lionychus quadrillum	A	2330	psamophile	Konst	•
Loricera pilicornis pilicornis	E				•
Masoreus wetterhallii wetterhallii	R	2330, 6110, 6190, 6210, 6240, 6250, 6260	psamophile	Dg	•
Microlestes corticalis	R				•
Microlestes fissuralis	A	40A0, 6110, 6190, 6210		Konst	•
Microlestes fulvibasis	R
Microlestes maurus maurus	E				•
Microlestes minutulus	E				•
Microlestes plagiatus	R				•
Microlestes schroederi	R	6210, 6240, 6250		Dg, Rar	•
Miscodera arctica	R	91T0, 91U0	psamophile	Dg, Rar	•
Molops piceus austriacus	A	9110, 9130, 9140, 9150, 9180, 9410		Konst, Rar	•
Nebria brevicollis	A				•
Nebria fuscipes	R
Nebria jockischii hoepfneri	A	3220/30/40	ripicole	Konst	•
Nebria livida livida	A		ripicole	Konst	•
Nebria picicornis picicornis	R	3220/30/40	ripicole	Dg, Rar	•
Nebria rufescens rufescens	R	3220/30/40, 6150	ripicole	Dg	•
Nebria salina	R				•
Nebria tatrica dumbirensis	R
Nebria tatrica fatrensis	R
Nebria tatrica komareki	R
Nebria tatrica tatrica	R
Notiophilus aestuans	E				•
Notiophilus aquaticus	A	4030, 4060, 4070/80, 5130, 6230, 6520, 91D0		Konst	•
Notiophilus biguttatus	A				•
Notiophilus germinyi	A	4030, 4060, 5130, 91T0, 91U0		Konst	•
Notiophilus laticollis	R	6110, 6190, 6210, 6240, 6250		Dg	•
Notiophilus palustris	E				•
Notiophilus rufipes	R	9170, 9190, 91G0, 91H0, 91I0		Dg	•
Ocys harpaloides	A
Ocys quinquestriatus quinquestriatus	E				•
Odacantha melanura	A				•
Olisthopus rotundatus rotundatus	A	4030, 5130		Konst	•
Olisthopus sturmii	R	40A0, 6110, 6190, 6210, 6240, 6250, 9170, 9190, 91G0, 91H0, 91I0		Dg	•
Omoglymmius germari	R				•
Omophron limbatum	A				•
Oodes gracilis	R	1340, 3130, 7210, 7230	subhalophile	Dg	•
Oodes helopioides	A				•
Ophonus ardosiacus	R	6210, 6240, 6250		Dg	•
Ophonus azureus	E				•
Ophonus brevicollis	A				•
Ophonus cordatus	R	6110, 6190, 6210		Dg	•
Ophonus cribricollis	R	6210, 6240, 6250		Dg, Rar	•
Ophonus diffinis	A	6210, 6240, 6250		Konst	•
Ophonus gammeli	R	9170, 9190, 91G0, 91H0, 91I0		Dg, Rar	•
Ophonus laticollis	A	6410, 6440, 6510		Konst	•
Ophonus melleti melleti	E				•
Ophonus parallelus	A				•
Ophonus puncticeps	E				•
Ophonus puncticollis	A	6110, 6190, 6210, 6240, 6250		Konst	•
Ophonus rufibarbis	E				•
Ophonus rupicola	E				•
Ophonus sabulicola	R	6210, 6240, 6250		Dg	•
Ophonus schaubergerianus	E				•
Ophonus stictus	A	6110, 6190, 6210, 6240, 6250		Konst	•
Ophonus subquadratus	A
Ophonus subsinuatus	A				•
Oreonebria castanea sumavica	R	8110, 8150, 8160, 8210, 8220, 8230, 9410		Dg, Rar	•
Oxypselaphus obscurus	A	91	hygrophile	Konst	•
Panagaeus bipustulatus	A	5130		Konst	•
Panagaeus cruxmajor	A	6410, 6440, 6510		Konst	•
Pangus scaritides	A
Paradromius linearis linearis	E				•
Paradromius longiceps	A	3130	hygrophile	Konst	•
Paradromius ruficollis	R	6150, 6230, 6520		Dg	•
Paranchus albipes	A				•
Paratachys bistriatus	A				•
Paratachys fulvicollis	A				•
Paratachys micros	R				•
Paratachys turkestanicus	A				•
Parazuphium chevrolatii praepannonicum	R
Parazuphium chevrolatii rebli	R	6210		Dg, Rar	•
Parophonus dejeani	A	6240, 6250		Konst, Rar	•
Parophonus hirsutulus	A	6240, 6250		Konst	•
Parophonus maculicornis	A				•
Parophonus mendax	A
Patrobus assimilis	R	7110, 7120, 7140, 7150, 91D0	tyrphophile, hygrophile	Dg	•
Patrobus atrorufus	A	91	hygrophile	Konst	•
Patrobus septentrionis australis	R	3130, 91E0, 91F0	hygrophile	Dg, Rar	•
Patrobus styriacus	R
Pedius longicollis	A	6240, 6250		Konst	•
Perigona nigriceps	E				•
Perileptus areolatus areolatus	R	3220/30/40	ripicole	Dg	•
Philorhizus crucifer crucifer	A	6240, 6250		Konst	•
Philorhizus notatus	A				•
Philorhizus quadrisignatus	A				•
Philorhizus sigma	A	3130	hygrophile	Konst	•
Platyderus rufus rufus	A	9170, 9190, 91G0, 91H0, 91I0		Konst	•
Platynus livens	R	3130, 91E0, 91F0	hygrophile	Dg, Rar	•
Poecilus cupreus cupreus	E				•
Poecilus koyi koyi	R	6210, 6240, 6250		Dg	•
Poecilus kugelanni	R				•
Poecilus lepidus lepidus	A				•
Poecilus puncticollis	R
Poecilus punctulatus	R	6210, 6240, 6250		Dg, Rar	•
Poecilus striatopunctatus	R
Poecilus szepligetti havelkai	R
Poecilus versicolor	E				•
Pogonus luridipennis	R	1340	halobiont	Dg, Rar	•
Pogonus peisonis	R	1340	halobiont	Dg, Rar	•
Polistichus connexus	R	6210, 6240, 6250		Dg, Rar	•
Porotachys bisulcatus	E				•
Pterostichus aethiops	A	9410		Konst	•
Pterostichus anthracinus anthracinus	A	3130	hygrophile	Konst	•
Pterostichus aterrimus aterrimus	R	3130	hygrophile	Dg, Rar	•
Pterostichus blandulus	R
Pterostichus burmeisteri burmeisteri	A	9110, 9130, 9140, 9150, 9180		Konst	•
Pterostichus cursor	R	1340	halophile	Dg, Rar	•
Pterostichus cylindricus	A	91E0, 91F0		Konst, Rar	•
Pterostichus diligens	A	91D0, 91E0	hygrophile	Konst	•
Pterostichus elongatus	R	3130	hygrophile	Dg, Rar	•
Pterostichus foveolatus	A				•
Pterostichus gracilis gracilis	R	3130	hygrophile	Dg, Rar	•
Pterostichus hungaricus	R
Pterostichus chameleon	R	3130	hygrophile	Dg, Rar	•
Pterostichus illigeri sudeticus	A	9410		Konst, Rar	•
Pterostichus incommodus	R	40A0, 9170, 9190, 91G0, 91H0, 91I0		Dg, Rar	•
Pterostichus inquinatus	R
Pterostichus latoricaensis	R	91	hygrophile	Dg, Rar	•
Pterostichus latoricaensis	R				•
Pterostichus leonisi	E				•
Pterostichus longicollis	A				•
Pterostichus macer macer	A	6240, 6250		Konst	•
Pterostichus madidus	R				•
Pterostichus melanarius melanarius	E				•
Pterostichus melas melas	A	40A0, 9170, 9190, 91G0, 91H0, 91I0		Konst	•
Pterostichus minor minor	A	3130	hygrophile	Konst	•
Pterostichus morio carpathicus	A
Pterostichus negligens negligens	R	6150, 8110, 8150, 8160, 8210, 8220, 8230		Dg	•
Pterostichus niger niger	A				•
Pterostichus nigrita	E				•
Pterostichus oblongopuntatus oblongopunctatus	A				•
Pterostichus ovoideus ovoideus	A	6410, 6440, 6510		Konst	•
Pterostichus pilosus pilosus	A	9410		Konst	•
Pterostichus pumilio pumilio	A	4070/80, 9410		Konst	•
Pterostichus quadrifoveolatus	A				•
Pterostichus rhaeticus	A	7110, 7120, 7140, 7150, 91D0, 91E0	hygrophile	Konst	•
Pterostichus rufitarsis cordatus	R	91D0, 9410		Dg	•
Pterostichus selmanni roubali	R	9410		Dg, Rar	•
Pterostichus strenuus	E				•
Pterostichus taksonyis	R	3130	hygrophile	Dg, Rar	•
Pterostichus tatricus	R
Pterostichus unctulatus	A	4070/80, 9410		Konst	•
Pterostichus vernalis	A				•
Rhysodes sulcatus	R	9110, 9130, 9140, 9150, 9180, 9170, 9190, 91G0, 91H0, 91I0, 9410		Dg, Rar	•
Scarites terricola	R
Sericoda quadripunctata	A				•
Sinechostictus decoratus	R				•
Sinechostictus doderoi	R	3220/30/40	ripicole	Dg, Rar	•
Sinechostictus millerianus	R	3220/30/40	ripicole	Dg, Rar	•
Sinechostictus ruficornis	R	3220/30/40	ripicole	Dg, Rar	•
Sinechostictus stomoides stomoides	R	3220/30/40	ripicole	Dg	•
Sphodrus leucophthalmus	A				•
Stenolophus discophorus	A	3270	ripicole	Konst, Rar	•
Stenolophus mixtus	A				•
Stenolophus skrimshiranus	A	3130	hygrophile	Konst	•
Stenolophus steveni	R	1340	halobiont	Dg, Rar	•
Stenolophus teutonus	E				•
Stomis pumicatus obenbergeri	A
Stomis pumicatus pumicatus	A				•
Syntomus foveatus	A	2330, 6260, 91T0, 91U0	psamophile	Konst	•
Syntomus obscuroguttatus	A	40A0, 91F0		Konst	•
Syntomus pallipes	A	40A0		Konst	•
Syntomus truncatellus	E				•
Synuchus vivalis vivalis	E				•
Tachys fulvicollis	A	1340	subhalophile	Konst, Rar	•
Tachys micros	R	3220/30/40	ripicole	Dg	•
Tachys turkestanicus	A		ripicole	Konst, Rar	•
Tachyta nana nana	A				•
Tachyura diabrachys	A				•
Tachyura haemorrhoidalis	A				•
Tachyura parvula	A				•
Tachyura parvula	A	3220/30/40	subripicole	Konst	•
Tachyura quadrisignata quadrisignata	A				•
Tachyura quadrisignata quadrisignata	A	3220/30/40	subripicole	Konst	•
Tachyura walkeriana	R				•
Thalassophilus longicornis	R	3220/30/40	ripicole	Dg, Rar	•
Trechoblemus micros	A				•
Trechus alpicola alpicola	R	9410		Dg	•
Trechus amplicollis	R	7110, 7120, 7140, 91D0, 9410	tyrphophile	Dg	•
Trechus austriacus	E				•
Trechus latus latus	A	9110, 9130, 9140, 9150, 9180	hygrophile	Konst	•
Trechus matejkai	R
Trechus montanellus	R	9410	hygrophile	Dg, Rar	•
Trechus obtusus obtusus	E				•
Trechus pilisensis pilisensis	A	9110, 9130, 9140, 9150, 9180, 91E0	hygrophile	Konst	•
Trechus pilisensis sudeticus	A	9110, 9130, 9140, 9150, 9180, 91E0	hygrophile	Konst	•
Trechus pulchellus pulchellus	A	9110, 9130, 9140, 9150, 9180, 91E0		Konst	•
Trechus pulpani	R
Trechus quadristriatus	E				•
Trechus rivularis	R				•
Trechus rivularis	R	7110, 7120, 7140, 7150, 91D0	hygrophile	Dg, Rar	•
Trechus rubem	A				•
Trechus secalis secalis	A				•
Trechus splendens	A	7110, 7120, 7140, 91D0, 91E0, 9410	hygrophile	Konst	•
Trechus striatulus	A	6150		Konst	•
Trechus tristis	A
Trichocellus cognatus	R				•
Trichocellus placidus	A				•
Trichotichnus laevicollis	A				•
Zabrus spinipes spinipes	R	6210		Dg, Rar	•
Zabrus tenebrioides tenebrioides	E				•

Abbreviation	Name	Description
R	relict species	These are species with the narrowest ecological valence, many of which are now relics. They are mostly rare and endangered species of natural, relatively undamaged ecosystems.
A	Adaptable species	These are species that inhabit more or less natural habitats or habitats close to their natural state. They also occur in secondary, well-regenerated biotopes, especially near their original areas.
E	Eurythopic species	These are species that often have no special requirements in terms of the character and quality of the environment, species of unstable, changing biotopes, as well as species inhabiting landscapes that are heavily anthropogenically influenced and damaged.

Abbreviation	Type	Description
Dg	Diagnostic	Species whose presence defines a given type of biotope complex in relation to other habitat types; in rare cases, a diagnostic species may also be a constant (or even dominant) species.
Rar	Rare	Species that are generally rare but occur more frequently within a given type of biotope complex.
Konst	constant	Species characteristic of a given type of biotope complex with regular occurrence in that type of biotope complex, but which are not exclusively linked to that type of biotope complex and also occur in other types of biotope complexes.

The Lexicon of Beetles of the Czech Republic